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The crew of the Serenity is getting desperate. They have lain lower and lower to protect their doctor, Simon Tam (Sean Maher), and his telepathic traumatized little sister, River (Summer Glau), from the alliance, whom he rescued her from a year earlier. This has made getting jobs harder and now they are desperate. When they take River on a robbery during which Reavers (humans who have gone crazy and turned into cannibals) attack, Simon decides that its time to leave the crew of the Serenity for his and River’s safety. She then mutters “Miranda” and goes berserk and nearly shoots Capt. Malcolm Reynolds (Nathan Fillion), until her brother says the safety word to put her to sleep. Reynolds decides to take them back on board for safety, only now an alliance operative (Chiwetel Ejiofor) is on their tracks and making Reynolds determined to find out what “Miranda” is and what the Alliance is hiding.

Sauron’s forces increase. His allies grow. The Ringwraiths return in an even more frightening form. Saruman’s army of Uruk Hai is ready to launch an assault against Aragorn and the people of Rohan. Yet, the Fellowship is broken and Boromir is dead. For the little hope that is left, Frodo and Sam march on into Mordor, unprotected. A number of new allies join with Aragorn, Gimli, Legolas, Pippin and Merry. And they must defend Rohan and attack Isengard. Yet, while all this is going on, Sauron’s troops mass toward the City of Gondor, for the War of the Ring is about to begin.

An ancient Ring thought lost for centuries has been found, and through a strange twist in fate has been given to a small Hobbit named Frodo. When Gandalf discovers the Ring is in fact the One Ring of the Dark Lord Sauron, Frodo must make an epic quest to the Cracks of Doom in order to destroy it! However he does not go alone. He is joined by Gandalf, Legolas the elf, Gimli the Dwarf, Aragorn, Boromir and his three Hobbit friends Merry, Pippin and Samwise. Through mountains, snow, darkness, forests, rivers and plains, facing evil and danger at every corner the Fellowship of the Ring must go. Their quest to destroy the One Ring is the only hope for the end of the Dark Lords reign!

While Frodo & Sam continue to approach Mount Doom to destroy the One Ring, unaware of the path Gollum is leading them, the former Fellowship aid Rohan & Gondor in a great battle in the Pelennor Fields, Minas Tirith and the Black Gates as Sauron wages his last war against Middle-Earth.